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² ˲ Ͳ ² ˲ : в ̳ - , . , 1622 2013 ...

-- [ 55 ] --

myrtifolium - . - , (Brundrett, 2009), . R. myrtifolium. R. myrtifolium . .

1. Brundrett M. Mycorrhizas in natural ecosystems. // Advances in ecological research. 2009. Vol. 21. P. 171-331.

2. Douglas, G.C., Heslin, M.C. and Reid, C. (1989) Isolation of Oidiodendron maius from Rhododendron and ultrastructural characterization of synthesized mycorrhizas. Canadian Journal of Botany, 67, 220612.

3. Hambleton S, Currah RS (1997). Fungal endophytes from the roots of alpine and boreal Ericaceae. Canadian Journal of Botany 75: 15701581.

4. Read DJ (1991) Mycorrhizas in ecosystems. Experientia 47: 376391.

5. Thormann, M.N.; Currah, R.S.; Bayley, S.E. 1999. The mycorrhizal status of the dominant vegetation along a peatland gradient in southern boreal Alberta, Canada.

Wetlands 19: 438-450.

6. ., . Rhododendron myrtifolium Scott et Kotschy . 䳿. 2011. . 5, 1, . 149-158.

7. .. / . // . , 1951. . 4. . 226-231.

8. .., .. . . : Գ, 2000, . 1. 365 .

9. , [ ] / .. , .. / , 2008. 105 .

10. .. / .. // .

, 1980. . 3-13.

11. .. // . , 1987.

12. . . : -, 2009. 912 .

13. .. . : . , 1976. 270 .

14. .. . . 1962.

360 581.9.524:502.7

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Shumska N.V. Ancient Beech Forests of Fageta sylvaticae subformation of the Halych National Nature Park: cenotical diversity. The results of ancient beech forests vegetation research of the Halych National Nature Park are presented. The forests are more than 100 years old. The characteristic of typical and unique phytocoenosis of Fageta sylvaticae subformation are presented. There phytocoenosis

belong to 11 associations. Five of them are included to Green Book of Ukraine:

Fagetum (sylvaticae) vincosum (minoris), Fagetum (sylvaticae) alliosum (ursini), Fagetum (sylvaticae) hederosum (helicis), Fagetum (sylvaticae) lunariosum (redivivae), Fagetum (sylvaticae) phyllitidosum (scolopendrii).

14684,8 2004 . ³ - , - , . , - [8].

11,9 . , 2,36 . ( 20 %). , Fageta sylvaticae 100 315,2 . arpineto betuli Fageta sylvaticae, arpineto betuli Querceto roboris Fageta sylvaticae, Tilio cordatae Carpineto betuli Querceto roboris-Fageta sylvaticae, Querceto roboris Fageta sylvaticae. Acer pseudoplatanus, Abies alba, Betula pendula. 3 2.

, , , , , 300 450 , ˳, , [6].

[1], -- - - .

, , , , - , , , .

Գ - [4, 9]. Fageta sylvaticae.

2008 2012 . 25 25 [3]. [7]. [5].

Fageta sylvaticae 110 , . ³ 100 120, 140 . 60 80 . Fagus sylvatica, Carpinus betulus, Tilia cordata, Acer pseudoplatanus.

100 28 32 36 (38) 42 (60) . 0,7 0,8; , , .

Fagetum (sylvaticae) caricosum (pilosae). F. sylvatica Carpinus betulus, Acer pseudoplatanus, Betula pendula. ϳ F. sylvatica, Tilia cordat, Acer pseudoplatanus. Euonymus verrucosa, Sambucus nigra, Daphne mezereum. , Carex pilosa, 30 80 %. Rubus hirtus (+ 20 %), Pulmonaria obscura, Paris quadrifolia, Dryopteris filix-mas, D.

carthusiana, Athyrium filix-femina, Galium odoratum, Stellaria holostea, Lamium maculatum, Lathyrus vernus, Milium effusum, Luzula pilosa, Polygonatum multiflorum, Galeobdolon luteum, Hepatica nobilis, Asarum europaeum . Anemone nemorosa (10 30 %), Isopyrum thalictroides, Scilla bifolia, Corydalis solida, Symphytum besseri, Galanthus nivalis.

362 Fagetum (sylvaticae) vincosum (minoris). 70 80 %. ϳ ; Tilia cordata, Acer platanoides, Acer pseudoplatanus. Vinca minor 50 100 %, Carex pilosa, Pulmonaria obscura, Paris quadrifolia, Dryopteris filixmas, D. carthusiana, Athyrium filix-femina, Galium odoratum, Stellaria holostea, Polygonatum multiflorum, Asarum europaeum.




140 .


Fagetum (sylvaticae) galiosum (odorati) . ϳ Fagus sylvatica (10 %), Tilia cordata (5 %), Acer psedoplatanus, Euonymus europaeus, Lonicera xylosteum, Daphne mezereum, Hedera helix. 60 80 %. Galium odoratum (30 60 %), Carex pilosa (+ 10 %), Aegopodium podagraria, Paris quadrifolia, Salvia glutinosa, Prenanthes purpurea, Circaea lutetiana, Pulmonaria obscura, Lilium martagon, Aposeris foetida, Sanicula europaea, Stellaria holostea, Lathyrus vernus, Galeobdolon luteum, Majanthemum bifolium, Asarum europaeum, Hepatica nobilis, Oxalis acetosella. Dryopteris filix-mas, D. carthusiana, Athyrium filixfemina, Gymnocarpium dryopteris, Dryopteris austriaca, Polystichum braunii, P. aculeatum.

Fagetum (sylvaticae) athyriosum (filix-feminae).

70 %. Euonymus verrucosa, Sambucus nigra. 60 80 % Athyrium filix-femina (30 60 %), Carex pilosa (+ 30 %), Dryopteris filix-mas, D. carthusiana, Pyrola rotundifolia, Galium odoratum, Asarum europaeum, Lamium maculatum, Polygonatum multiflorum, Lathyrus vernus.

Fagetum (sylvaticae) nudum . 80 %. ϳ Sorbus aucuparia, Lonicera xylosteum, Fagus sylvatica, Acer pseudoplatanus, Abies alba, Picea abies. 1 5 % Athyrium filix-femina, Dryopteris carthusiana, Prenanthes purpurea, Luzula pilosa, Huperzia selago, Majanthemum bifolium, Oxalis acetosella .

, , Fagetum (sylvaticae) dentariosum (glandulosae) 60 100 %. . Dentaria glandulosa (50 80 %), Anemone nemorosa, Pulmonaria obscura, Leucojum vernum, Gagea lutea, Scilla bifolia, Chrysosplenium alternifolium, Symphytum cordatum, Adoxa moschatellina, Corydalis solida. , Aegopodium podagraria (20 30), Dryopteris filix-mas, D. carthusiana, Athyrium filix-femina .

Fagetum (sylvaticae) alliosum (ursini) , .

70 %. ϳ Sambucus nigra, Euonymus europaea, Carpinus betulus, Fagus sylvatica, Tilia cordata, Acer pseudoplatanus. 80 100 % Allium ursinum (50 90 %). Aegopodium podagraria (20 30 %), Asarum europaeum, Galium odoratum, Dentaria bulbifera, Glechoma hederacea, Dryopteris filix-mas, D. carthusiana, Athyrium filix-femina, Aposeris foetida, Pulmonaria obscura, Symphytum cordatum, Ficaria verna (30 60 %), Scilla bifolia, Galanthus nivalis, Gagea lutea.

Fagetum (sylvaticae) aegopodiosum (podagrariae), Aegopodium podagraria (50 80 %) Allium ursinum.

- Fagetum (sylvaticae) hederosum (helicis) Hedera helix 30 60 %. Sambucus nigra, Euonymus verrucosa. , Mercurialis perennis (+ 10 %), Phyllitis scolopendrium (+ 10 %), Polypodium vulgare (+ 20 %), Campanula rapunculoides, Asarum europaeum, Asplenium trichomanes.

Fagetum (sylvaticae) lunariosum (redivivae) , . 70 80 %. Sambucus nigra, Acer pseudoplatanus Euonymus verrucosa; Fagus sylvatica.

60 80 %, Lunaria rediviva 40 60 %. Mercurialis perennis, Stachys sylvatica, Aegopodium podagraria, Symphytum cordatum, Lamium maculatum, Asarum europaeum; - Phyllitis scolopendrium (10 %), Polypodium vulgare (+ 10 %), Mercurialis perennis, Geranium robertianum, Chelidonium majus, Asarum europaeum.

, - , Fagetum (sylvaticae) phyllitidosum (scolopendrii). ϳ Sambucus nigra, Fagus sylvatica. 30 60 %, Phyllitis scolopendrium 30 50 %. Asarum europaeum 364 (10 %), Polypodium vulgare (10 %), Geranium robertianum, Polystichum braunii.

, Fageta sylvaticae , . 11 , [2] Fagetum (sylvaticae) vincosum (minoris), Fagetum (sylvaticae) alliosum (ursini), Fagetum (sylvaticae) hederosum (helicis), Fagetum (sylvaticae) lunariosum (redivivae), Fagetum (sylvaticae) phyllitidosum (scolopendrii).

1. / . . .. . :

. , 1977. 304 .

2. / . . .. ij .: , 2009. 448 .



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