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, , , , , , , . () , . . (Shugart, 1999, , 2011) ( ), , 280 1 50 (, 2011).

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1. .. // . . , . , 2012. . 99-103.

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59. . 4-6.

3. .. Soldanella hungarica Simonk.

(Primulaceae) // . 2007. . 64, 5. . 651-659.

282

4. .., .. // . 2010. 2(6). . 46-49.

5. .. : -. , 2003. 244 .

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. . - . . : . 2009. 1-2 (39). . 113-123.

7. г .., г .. // ( ). . . 14-18 2002 . . 2002. 2. . 455-459.

8. г .., .. - () // . . 2008. 24. . 24-34.

9. г .., .., Lachat T., .., г .., ..

- (Coleoptera, Carabidae) ( ) // . . 2011. .

31. . 16-28.

10. . // ³ . . 2003. . 33. . 60-64.

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// . . 2007. 18. . 72-82.

12. Astous. D., Villard M.-A. Effects of Selection Harvesting on Bark Invertebrates and Nest Provisioning Rate in an Old Forest Specialist, the Brown Creeper (Certhia americana) // Ecoscience. 2012. Vol. 19 (2). P. 106-112.

13. Baker J., French K., Whelan R.J. The edge effect and ecotonal species: bird communities across a natural edge in southeastern Australia // Ecology. 2002. Vol. 83. P. 3048-3059.




140 .


14. Imbeau L., Drapeau P., Mnkknen M. Are forest birds categorised as edge species strictly associated with edges? // Ecography. 2003. Vol. 26, Iss. 4. P.

514-520.

15. Manolis J. C., Andersen D. E., Cuthbert F. J. Edge effect on nesting success of ground nesting birds near regenerating clearcuts in a forest-dominated landscape // The Auk. 2002. Vol. 119 (4). P. 955-970.

16. Poulin J.-F., Villard M.-A., Hach S. Short-Term Demographic Response of an Old Forest Specialist to Experimental Selection Harvesting // Ecoscience. 2010. Vol. 17 (1). P. 20-27.

17. Ries L., Fletcher R. J., Battin J., Sisk T. D. Ecological responses to habitat edges:

mechanisms, models, and variability explained // Annual Review of Ecology, Evolution, and Systematics. 2004. Vol. 35. P. 491-522.

18. Walker S., Wilson J. B., Steel J. B., Rapson G. L., Smith B., King W. McG., Cottam Y. H. Properties of ecotones: Evidence from five ecotones objectively determined from a coastal vegetation gradient // Journal of Vegetation Science. 2003. Vol. 14, Iss. 4. P. 579-590.

595.762.12(477.87):502.7 - (COLEOPTERA, CARABIDAE) ˲Ѳ ² ХȻ

   

, . , , . , Rizun V.B., Slobodyan O.M. Ground-beetles (Coleoptera, Carabidae) of virgin forests of the Nature Reserve Gorgany. Species and dominance structure of carabid communities in the beech-fir-spruce, spruce and cedar-spruce virgin forests are given. The fifteen Carabidae species have been identified. Within 16 Carpathian endemic species only 3 species (Carabus obsoletus Strm., Strm., Pterostichus foveolatus (Duft.), Pterostichus pilosus (Host)) were found.

() 1996 . 800-1754 . . . : () , ( ) . ˳ 84,4 %, () 12,2 %, , 3,4 % . 900-1400 . .

- .. г (2002, 2003), .. (2004) 116 (г, 2006).

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- (Bombina variegata (L.)) (г, , г, 2006).

. - (-1, -2, -3), 11 , .

-1 26, 1000-1040 , 180-1900, 15-200, ;

: N4847'490''; E2430'619'' 3 . - -⳿ . : 442,., (): 0,7-0,9; 3.

284 (Picea abies (L.) Karst): 138 , 11 , 8 , 158 ; : 80-140 .; ; ; : 44,0 , 25,3 ; :

105,0 , 31,9 . (Abies alba Mill.): 150 , 6 , , 157 ; : 80-140 .; ; ; : 46,8 , 20,9 .; : 116,7 , 29,1 . (Fagus sylvatica (Bess.) Reichb.): 94 ,

, _ , _ 96 ; : 80-120 .; ; ; :

32,0 , 23,9 .; : 78,3 , 30,1 . (Acer pseudoplatanus L.): 10 ;

: ~ 120-150 ; ; ; : 33,9 , 23,4 .; :

83,0 , 45,5 .

ϳ. : 4321,.. dz: 1 1520 . 260 . . . : 5,0 , 0,60 . : 85 . : .

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: 82,.,; 0,7-0,9; 0,86; , .



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